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Research article - Peer-reviewed, 2021

Mixing effects on Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) productivity along a climatic gradient across Europe

Ruiz-Peinado, Ricardo; Pretzsch, Hans; Lof, Magnus; Heym, Michael; Bielak, Kamil; Aldea, Jorge; Barbeito, Ignacio; Brazaitis, Gediminas; Drossler, Lars; Godvod, Ksistof; Granhus, Aksel; Holm, Stig-Olof; Jansons, Aris; Makrickiene, Ekaterina; Metslaid, Marek; Metslaid, Sandra; Nothdurft, Arne; Reventlow, Ditlev Otto Juel; Sitko, Roman; Stankeviciene, Gintare;
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Mixed-species stands have been found to be more productive than would be expected from the performance of their component species in monocultures due to facilitation and complementarity between species, although these interactions depend on the combination of species present. Our study focuses on monospecific and mixed species stands of Scots pine and Norway spruce using 20 triplets established in nine countries along a climatic gradient across Europe. Differences in mean tree and stand characteristics, productivity and stand structure were assessed. Basal area increment in mixed stands was 8% higher than expected while volume increment was only 2% greater. Scots pine trees growing in mixed-species stands showed 11% larger quadratic mean diameter, 7% larger dominant diameter, 17% higher basal area and 25% higher stand volume than trees growing in mono specific stands. Norway spruce showed only a non-significant tendency to lower mean values of diameters, heights, basal area, as well standing volume in mixtures than monocultures. Stand structure indices differed between mixed stands and monocultures of Scots pine showing a greater stratification in mixed-species stands. Furthermore, the studied morphological traits showed little variability for trees growing in monospecific stands, except for diameter at breast height, crown length and crown length ratio. For trees growing in mixed stands, all the morphological traits of the trees were identified as different. Some of these morphological traits were associated with relative productivity. Nevertheless, relative productivity in mixed-species stands was not related to site conditions.


Monospecific and mixed-species stands; Overyielding; Stand structure; Complementarity; Triplet approach

Published in

Forest Ecology and Management
2021, volume: 482, article number: 118834
Publisher: ELSEVIER

Authors' information

Ruiz-Peinado, Ricardo
Instituto Nacional Investigacion Tecnologia Agraria Alimentaria (INIA)
Pretzsch, Hans
Technical University of Munich
Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre
Heym, Michael
Technical University of Munich
Bielak, Kamil
Warsaw University of Life Sciences
Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre
Barbeito, Ignacio
Swedish University of Agricultural Sciences, Southern Swedish Forest Research Centre
Brazaitis, Gediminas
Vytautas Magnus University
Drössler, Lars
Ilia State University
Godvod, Ksistof
Vytautas Magnus University
Granhus, Aksel
Norwegian Institute of Bioeconomy Research (NIBIO)
Holm, Stig-Olof
Umeå University
Jansons, Aris
Latvian State Forest Research Institute Silava
Makrickiene, Ekaterina
Vytautas Magnus University
Metslaid, Marek
Estonian University of Life Sciences
Metslaid, Sandra
Estonian University of Life Sciences
Nothdurft, Arne
University of Bodenkultur Wien
Reventlow, Ditlev Otto Juel
University of Copenhagen
Sitko, Roman
Tech Univ Zvolen TUZVO
Stankeviciene, Gintare
Vytautas Magnus University
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UKÄ Subject classification

Forest Science

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